In Melbourne's first home match of the 2014 season at AAMI Park, Smith will break Matt Geyer's club games record of 262 first-grade games while Slater will play his 250th match.
Storm enter the Round 2 clash against the Panthers with plenty of momentum after a spectacular comeback victory in their season opener against Manly last weekend. In addition, Melbourne Captain Cameron Smith ended speculation on his playing future during the week, committing to the club for a further four seasons much to the delight of the Storm faithful who will continue to see their skipper run out onto AAMI Park for many years to come.
Speaking to the media on Tuesday morning, Melbourne fullback Billy Slater acknowledged the enormity of Storm's return to AAMI Park in 2014.
"It's a big week for the club, a big week with re-signing Cam, playing my 250th game, our first game at AAMI Park for the year and capped off with Cam breaking the all time games record of Matt Geyer's 262 games," Slater said.
To celebrate Smith and Slater's milestones, Melbourne Storm fans will get the chance to support their heroes with face masks provided by Programmed.
In other news, Storm halfback and 2013 Dally M Medallist Cooper Cronk will line-up for his first match of the 2014 season after being named in Craig Bellamy's squad to take on Penrith. For a full list of team news, click here.